skip to Main Content

MSc. Danny Haelewaters, Ph.D.

Research focus:

– natural enemies of ladybirds (Coleoptera: Coccinellidae)
– biology and patterns of speciation of ectoparasitic fungi (Ascomycota: Laboulbeniomycetes)
– evolutionary biology of ascomycetous fungi
– species diversity and evolution of microfungi
– invasion ecology


twitter: @dhaelewa



– 2012 – 2018: Ph.D., Department of Organismic and Evolutionary Biology, Harvard University, USA. Thesis: Studies of the Laboulbeniomycetes: Diversity, evolution, and patterns of speciation. Research advisor: Donald H. Pfister.
– 2009 – 2012: MSc., Ghent University, Faculty of Sciences, Belgium. Thesis: Laboulbeniales—Exploring and testing DNA extraction protocols, carrion beetle hosts and species in ‘De Kaaistoep’ (The Netherlands). Research advisors: Annemieke Verbeken, André De Kesel.
– 2006 – 2009: Bachelor of Science, Faculty of Sciences, Ghent University, Belgium.
– 2001 – 2004: Bachelor in Veterinary Sciences, Faculty of Pharmaceutical, Biomedical and Veterinary Sciences, University of Antwerp, Belgium.

Professional appointments:

– Sep 2021 – present: Researcher, Institute of Entomology, Biology Centre, Czech Academy of Sciences, Czech Republic.
– Nov 2020 – present: Junior Postdoctoral Fellow, Department of Biology, Ghent University, Belgium.
– 2019 – present: Researcher, Department of Zoology, Faculty of Science, University of South Bohemia, Czech Republic.
– 2018 – 2020: Postdoctoral Research Assistant, Department of Botany and Plant Pathology, Purdue University, USA.
– 2018 – 2019: Postdoctoral Researcher, Faculty of Science, University of South Bohemia, Czech Republic.
– 2017 (summer): Short-Term Research Fellow, Smithsonian Tropical Research Institute, Panama.
– 2016 – present: Associate Researcher, Herbario UCH, Universidad Autónoma de Chiriquí, Panama.

Internships and field experience:

– 2021: Croatia: Krka National Park
– 2019: Honduras: Cusuco National Park
– 2015 – 2019: Panama: Chiriquí (Boquete, David, Parque Nacional Volcán Barú, Potrerillos Abajo, Reserve Forestal de Fortuna, Volcán), Colón (Gamboa, Parque Nacional Soberanía), Darién (Chucanti Nature Reserve), Panamá Oeste (Barro Colorado Island)
– 2014: Peninsular Malaysia, 2 weeks.
– 2012 – 2018: Boston Harbor Islands National Recreation Area, Massachusetts, USA.
– 2012: Costa Rica, 1 month.
– 2009: Netherlands Forensic Institute, Den Haag, The Netherlands, 3 months.

Editorial positions:

– 2020 – present: Associate Editor, IMA Fungus
– 2020 – present: Member of Board of Editors, Neotropical Naturalist
– 2020 – present: Subject Editor, Biodiversity Data Journal
– 2020: Guest Editor, Mycologia 112(6) (Special Issue: Animal-Associated Fungi)
– 2019 – present: Member of Editorial Board, Parasite
– 2019 – present: Member of Editorial Board, Sydowia
– 2019 – present: Subject Editor, MycoKeys
– 2018 – 2021: Special Issue Editor, Northeastern Naturalist 25(Special Issue 9) (Boston Harbor Islands National Recreation Area: Overview of Recent Research)
– 2018 – 2019: Editor, IOBC-WPRS Bulletin volumes 137, 145


– Fall 2021: Lecturer in Doctoral Schools Natural Sciences, (Bioscience) Engineering, and Life Sciences and Medicine (X000947: Writing Academic Papers).
– Summer 2017: Universidad Autónoma de Chiriquí, Panama. Instructor of “DNA extraction techniques for Laboulbeniales fungi” workshop, in Spanish. Co-instructed with Tina A. Hofmann.
– 2015 – 2017: Harvard University, USA. Teaching Fellow in Organismic and Evolutionary Biology (OEB54: Biology of the Fungi). Taught with Donald H. Pfister. Recipient of Certificate of Distinction for Teaching Excellence.
– 2015 – 2016: Teaching Fellow in Organismic and Evolutionary Biology (OEB59: Plants and Human Affairs). Taught with Donald H. Pfister.
– 2013 – 2015: Teaching Fellow in Molecular and Cellular Biology (LS1b: Genetics, Genomics, and Evolution). Taught with Kevin Eggan, Hopi Hoekstra, Maryellen Ruvolo, Hopi Hoekstra, and Pardis Sabeti.


Bachelor’s theses:
Verdegem, S. (2021) DNA barcoding of Fungi collected at the Boston Harbor Islands, Massachusetts (Ghent University, Belgium)
Van Caenegem, W. (2021) Hyperparasitic interactions of Laboulbeniales on bat flies in Trinidad and Panama and the description of two new species of Gloeandromyces (Ghent University, Belgium)
Mironova, P. (2020) Een morfologische studie van Arthrorhynchus nycteribiae (Laboulbeniales) (Ghent University, Belgium)
Wang, T.W. (2016) Herpomyces: phylogeny, taxonomy, and ecology of a fungal genus parasitic on cockroaches (Harvard University, USA)

Master’s theses:
Santamaria, B.M. (started 2021) How do biotic and abiotic factors affect parasitism? A case study with ectoparasitic Laboulbeniales fungi (Ghent University, Belgium)
Suvorova, G. (started 2020) Effects of biotic and abiotic traits on a hyperparasite system of bats, bat flies and Laboulbeniales fungi in the tropics (Erasmus Mundus Masters Course in Tropical Biodiversity and Ecosystem)
Dumolein, I. (2021) The bat as a house for different residents: a tritrophic survey of bats, flies, and fungi in a Honduran cloud forest (Ghent University, Belgium)
Lubbers, M (2021) Peculiar parasites: on the ecology and genetics of Laboulbeniales (Leiden University, The Netherlands)
de Groot, M.D. (2021) Laboulbeniales hyperparasites of bat flies: distributions, host specificity, and abiotic traits (Leiden University, The Netherlands)
De Weggheleire, S. (2019) Biodiversity of the genus Laboulbenia: are there cryptic species? (University of Antwerp, Belgium)

Committee member for Ph.D. students:
Law, H. (started 2022) (Purdue University, USA)
Musa, N.N. (started 2021) Diversity and interaction study on Coccinellidae (Coleoptera) from several different landscapes in Peninsular Malaysia (Universiti Kebangsaan Malaysia)
Stallman, J.K. (started 2020) Leveraging tropical collections and molecular data to understand the biodiversity and evolutionary history of the Leotiomycetes (Ascomycota) (Purdue University, USA)
Schoutteten, N. (started 2019) A hidden and sneaky strategy: mechanisms and importance of mycoparasitism in the evolution of Basidiomycota (Ghent University, Belgium)


Google Scholar; ORCIDResearchGate

Research papers affiliated to University of South Bohemia:

Haelewaters D., Hofmann T.A., Romero-Olivares A.L., 2021: Ten simple rules for Global North researchers to stop perpetuating helicopter research in the Global South. PLoS Computational Biology 17 (8): e1009277.

Crous P.W., Lombard L., Sandoval-Denis M., Seifert K.A., Schroers H.J., Chaverri P., Gené J., Guarro J., Hirooka Y., Bensch K., Kema G.H.J., Lamprecht S.C., Cai L., Rossman A.Y., Stadler M., Summerbell R.C., Taylor J.W., Ploch S., Visagie C.M., Yilmaz N., Frisvad J.C., Abdel-Azeem A.M., Abdollahzadeh J., Abdolrasouli A., Akulov A., Alberts J.F., Araújo J.P.M., Ariyawansa H.A., Bakhshi M., Bendiksby M., Ben Hadj Amor A., Bezerra J.D.P., Boekhout T., Câmara M.P.S., Carbia M., Cardinali G., Castañeda-Ruiz R.F., Celis A., Chaturvedi V., Collemare J., Croll D., Damm U., Decock C.A., de Vries R.P., Ezekiel C.N., Fan X.L., Fernández N.B., Gaya E., González C.D., Gramaje D., Groenewald J.Z., Grube M., Guevara-Suarez M., Gupta V.K., Guarnaccia V., Haddaji A., Hagen F., Haelewaters D., Hansen K., Hashimoto A., Hernández-Restrepo M., Houbraken J., Hubka V., Hyde K.D., Iturriaga T., Jeewon R., Johnston P.R., Jurjević Ž., Karalti İ., Korsten L., Kuramae E.E., Kušan I., Labuda R., Lawrence D.P., Lee H.B., Lechat C.L.L., Li H.Y., Litovka Y.A., Maharachchikumbura S.S.N., Marin-Felix Y., Kemkuignou B.M., Matočec N., McTaggart A.R., Mlčoch P., Mugnai L., Nakashima C., Nilsson R.H., Noumeur S.R., Pavlov I.N., Peralta M.P., Phillips A.J.L., Pitt J.I., Polizzi G., Quaedvlieg W., Rajeshkumar K.C., Restrepo S., Rhaiem A., Robert J., Robert V., Rodrigues A.M., Salgado-Salazar C., Samson R.A., Santos A.C.S., Shivas R.G., Souza-Motta C.M., Sun G.Y., Swart W.J., Szoke S., Tan Y.P., Taylor J.E., Taylor P.W.J., Tiago P.V., Váczy K.Z., van de Wiele N., van der Merwe N.A., Verkley G.J.M., Vieira W.A.S., Vizzini A., Weir B.S., Wijayawardene N.N., Xia J.W., Yañez-Morales M.J., Yurkov A., Zamora J.C., Zare R., Zhang C.L., Thines M., 2021: Fusarium: more than a node or a foot-shaped basal cell. Studies in Mycology 98: 100116.

Gonçalves S.C., Haelewaters D., Furci G., Mueller G.M., 2021: Include all fungi in biodiversity goals. Science 373 (6553): 403.

Koukol O., Haelewaters D., 2021: Laboulbeniales – nejprapodivnější houboví paraziti / Laboulbeniales – the most peculiar fungal parasites. Časopis Živa, ročník 1/2021: 8–11.

Martin T.E., Jones S.E.I., Creedy T.J., Hoskins H.M.J., McCann N.P., Batke S.P., Kelly D.L., Kolby J.E., Downing R., Zelaya S.M.S., Green S.E.W., Lonsdale G., Brown T., Waters S., Rodríguez-Vásquez F., McCravy K.W., D’Souza M.L., Crace D., Nuñez-Mino J.M., Haelewaters D., Medina-van Berkum P., Phipps C.D., Barker R.J., Castañeda F., Reid N., Jocqué M., 2021: A review of the ecological value of Cusuco National Park: an urgent call for conservation action in a highly threatened Mesoamerican cloud forest. Journal of Mesoamerican Biology 1 (1): 6–50.

Medina-van Berkum P., Haelewaters D., Thompson J.R.E., Uribe E., Vulinec K., Martin T.E., 2021: Bat assemblages and their ectoparasites in a Honduran cloud forest: Effects of disturbance and altitude. Journal of Mesoamerican Biology 1 (1): 51–69.

Haelewaters D., Schoutteten N., Medina-van Berkum P., Martin T.E., Verbeken A., Aime M.C., 2021: Pioneering a fungal inventory at Cusuco National Park, Honduras. Journal of Mesoamerican Biology 1 (1): 111–131.

Dima B., Brandrud T.E., Corriol G., Jansen G.M., Jordal J.B., Khalid A.N., Larsson E., Lorås J., Morozova O.V., Naseer A., Noordeloos M.E., Rossi W., Santamaria S., Sarwar S., Sesli E., Usman M., Afshan N.S., Ahmad I., Banerjee A., Banerjee K., Bendiksen E., Colombo D.R.S., De Kesel A., Dovana F., Ferisin G., Hussain S., Islam S., Jesus A.L., Kaygusuz O., Krisai-Greilhuber I., Mahammad S., Mishra D.K., Nath P.S., da Paixão S.C.O., Panja B., Papp V., Pires-Zottarelli C.L.A., Radnóti Á., Rana D., Saha R., Türkekul İ., Haelewaters D., 2021: Fungal Systematics and Evolution: FUSE 7. Sydowia 73: 271–340.

Biranvand A., Nedvěd O., Nattier R., Nepaeva E., Haelewaters D., 2021: Review of the genus Hippodamia (Coleoptera: Coccinellidae) in the Palearctic region. Oriental Insects 55 (2): 293–304.

Mešić A., Haelewaters D., Tkalčec Z., Liu J., Kušan I., Aime M.C., Pošta A., 2021: Inocybe brijunica sp. nov., a new ectomycorrhizal fungus from Mediterranean Croatia revealed by morphology and multilocus phylogenetic analysis. Journal of Fungi 7 (3): 199.

Sulistyo B.P., Ryberg M., Haelewaters D., Larsson K.H., 2021: Multigene phylogeny and taxonomic revision of Atheliales s.l.: reinstatement of three families and one new family, Lobuliciaceae fam. nov. Fungal Biology 125 (3): 239–255.

Gómez-Zapata P.A., Haelewaters D., Quijada L., Pfister D.H., Aime M.C., 2021: Notes on Trochila (Ascomycota, Leotiomycetes), with new species and combinations. MycoKeys 78: 21–47.

Perreau M., Haelewaters D., Tafforeau P., 2021: A parasitic coevolution since the Miocene revealed by propagation phase-contrast synchrotron X-ray microtomography and the study of natural history collections. Scientific Reports 11 (1): 2672.

Haelewaters D., Blackwell M., Pfister D.H., 2021: Laboulbeniomycetes: Intimate fungal associates of arthropods. Annual Review of Entomology 66: 257–276.

Blackwell M., Haelewaters D., Pfister D.H., 2020: Laboulbeniomycetes: Evolution, natural history, and Thaxter’s final word. Mycologia 112 (6): 1048–1059.

Haelewaters D., Kasson M.T., 2020: Animal-Associated Fungi: Editorial. Mycologia 112 (6): 1045–1047.

Haelewaters D., Hiller T., Kemp E.A., van Wielink P.S., Shapiro-Ilan D.I., Aime M.C., Nedvěd O., Pfister D.H., Cottrell T.E., 2020: Mortality of native and invasive ladybirds co-infected by ectoparasitic and entomopathogenic fungi. PeerJ 8: e10110.

Haelewaters D., Dima B., Abdel-Hafiz B.I.I., Abdel-Wahab M.A., Abul-Ezz S.R., Acar I., Aguirre-Acosta E., Aime M.C., Aldemir S., Ali M., Ayala-Vásquez O., Bakhit M.S., Bashir H., Battistin E., Bendiksen E., Castro-Rivera R., Çolak Ö.F., De Kesel A., de la Fuente J.I., Dizkırıcı A., Hussain S., Jansen G.M., Kaygusuz O., Khalid A.N., Khan J., Kiyashko A.A., Larsson E., Martínez-González C.R., Morozova O.V., Niazi A.R., Noordeloos M.E., Pham T.H.G., Popov E.S., Psurtseva N.V., Schoutteten N., Sher H., Türkekul İ., Verbeken A., Ahmad H., Afshan N.S., Christe P., Fiaz M., Glaizot O., Liu J., Majeed J., Markotter W., Nagy A., Nawaz H., Papp V., Péter Á., Pfliegler W.P., Qasim T., Riaz M., Sándor A.D., Szentiványi T., Voglmayr H., Yousaf N., Krisai-Greilhuber I., 2020: Fungal Systematics and Evolution: FUSE 6. Sydowia 72: 231–356.

Wijayawardene N.N., Hyde K.D., Dai D.Q., Tang L.Z., Aptroot A., Castañeda-Ruiz R.F., Druzhinina I.S., Cai F., Ekanayaka A.H., Erdoğdu M., Fiuza P.O., Gentekaki E., Goto B.T., Haelewaters D., Hongsanan S., Jeewon R., Kirk P.M., Jayalal R.G.U., Karunarathna S.C., Wanasinghe D.N., Lumbsch H.T., Madrid H., Maharachchikumbura S.S.N., Monteiro J.S., Shivaprakash N., Pfliegler W.P., Phillips A.J.L., Saxena R.K., Stadler M., Tian Q., Tokarev Y.S., Tsurykau A., Ertz D., Lee H.B., Etayo J., Vizzini A., Jones E.B.G., Lin C.G., Li W.J., Dai Y.C., Fan X.L., McKenzie E.H.C., Shivas R.G., Hustad V., Leontyev D.V., de Hoog G.S., Niskanen T., Boekhout T., Gaya E., Thines M., 2020: A dynamic portal for a community-driven, continuously updated classification of Fungi and fungus-like organisms: Mycosphere 11 (1): 1514–1526.

Haelewaters D., De Kesel A., 2020: Checklist of thallus-forming Laboulbeniomycetes from Belgium and the Netherlands, including Hesperomyces halyziae and Laboulbenia quarantenae spp. nov. MycoKeys 71: 23–86.

Saba M., Haelewaters D., Pfister D.H., Khalid A.N., 2020: New species of Pseudosperma (Agaricales, Inocybaceae) from pine-dominated forests of western Himalayas. Mycokeys 69: 1–31.

Wijayawardene N.N., Hyde K.D., Al-Ani L.K.T., Tedersoo L., Haelewaters D., Rajeshkumar K.C., Zhao R.L., Aptroot A., Leontyev D.V., Saxena R.K., Tokarev Y.S., Dai D.Q., Letcher P.M., Stephenson S.L., Ertz D., Lumbsch H.T., Kukwa M., Issi I.V., Madrid H., Phillips A.J.L., Selbmann L., Pfliegler W.P., Horváth E., Bensch K., Kirk P.M., Kolaříková K., Raja H.A., Radek R., Papp V., Dima B., Ma J., Malosso E., Takamatsu S., Rambold G., Gannibal P.B., Triebel D., Gautam A.K., Avasthi S., Suetrong S., Timdal E., Fryar S.C., Delgado G., Réblová M., Doilom M., Dolatabadi S., Pawłowska J., Humber R.A., Kodsueb R., Sánchez-Castro I., Goto B.T., Silva D.K.A., de Souza F.A., Oehl F., da Silva G.A., Silva I.R., Błaszkowski J., Jobim K., Maia L.C., Barbosa F.R., Fiuza P.O., Divakar P.K., Shenoy B.D., Castañeda-Ruiz R.F., Somrithipol S., Lateef A.A., Karunarathna S.C., Tibpromma S., Mortimer P.E., Wanasinghe D.N., Phookamsak R., Xu J., Wang Y., Tian F., Alvarado P., Li D.W., Kušan I., Matočec N., Maharachchikumbura S.S.N., Papizadeh M., Heredia G., Wartchow F., Bakhshi M., Boehm E., Youssef N., Hustad V.P., Lawrey J.D., Santiago A.L.C.M.A., Bezerra J.D.P., Souza-Motta C.M., Firmino A.L., Tian Q., Houbraken J., Hongsanan S., Tanaka K., Dissanayake A.J., Monteiro J.S., Grossart H.P., Suija A., Weerakoon G., Etayo J., Tsurykau A., Vázquez V., Mungai P., Damm U., Li Q.R., Zhang H., Boonmee S., Lu Y.Z., Becerra A.G., Kendrick B., Brearley F.Q., Motiejūnaitė J., Sharma B., Khare R., Gaikwad S., Wijesundara D.S.A., Tang L.Z., He M.Q., Flakus A., Rodriguez-Flakus P., Zhurbenko M.P., McKenzie E.H.C., Stadler M., Bhat D.J., Liu J.K., Raza M., Jeewon R., Nassonova E.S., Prieto M., Jayalal R.G.U., Erdoğdu M., Yurkov A., Schnittler M., Shchepin O.N., Novozhilov Y.K., Silva-Filho A.G.S., Liu P., Cavender J.C., Kang Y., Mohammad S., Zhang L.F., Xu R.F., Li Y.M., Dayarathne M.C., Ekanayaka A.H., Wen T.C., Deng C.Y., Pereira O.L., Navathe S., Hawksworth D.L., Fan X.L., Dissanayake L.S., Kuhnert E., Grossart H.P., Thines M., 2020: Outline of Fungi and fungus-like taxa. Mycosphere 11 (1): 1060–1456.

De Kesel A., Gerstmans C., Haelewaters D., 2020: Catalogue of the Laboulbeniomycetes of Belgium. Sterbeeckia 36: 3–143.

Kiran M., Sattar A., Zamir K., Haelewaters D., Khalid A.N., 2020: Additions to the genus Chroogomphus (Boletales, Gomphidiaceae) from Pakistan. MycoKeys 66: 23–38.

Hyde K.D., Dong Y., Phookamsak R., Jeewon R., Bhat D.J., Jones E.B.G., Liu N., Abeywickrama P.D., Mapook A., Wei D., Perera R.H., Manawasinghe I.S., Pem D., Bundhun D., Karunarathna A., Ekanayaka A.H., Bao D.F., Li J., Samarakoon M.C., Chaiwan N., Lin C.G., Phutthacharoen K., Zhang S.N., Senanayake I.C., Goonasekara I.D., Thambugala K.M., Phukhamsakda C., Tennakoon D.S., Jiang H.B., Yang J., Zeng M., Huanraluek N., Wijesinghe S.N., Tian Q., Tibpromma S., Brahmanage R.S., Boonmee S., Huang S.K., Thiyagaraja V., Lu Y.Z., Jayawardena R.S., Dong W., Yang E.F., Singh S.K., Singh S.M., Rana S., Lad S.S., Anand G., Devadatha B., Niranjan M., Sarma V.V., Liimatainen K., Niskanen T., Overall A., Alvarenga R.L.M., Gibertoni T.B., Pfliegler W.P., Horváth E., Imre A., Alves A.L., da Silva Santos A.C., Tiago P.V., Bulgakov T.S., Wanasinghe D.N., Liu J.K., Doilom M., Maharachchikumbura S.S.N., Rajeshkumar K.C., Haelewaters D., Mortimer P.E., Zhao Q., Lumyong S., Xu J., Sheng J., 2020: Fungal diversity notes 1151–1275: taxonomic and phylogenetic contributions on genera and species of fungal taxa. Fungal Diversity 100 (1): 5–277.

He M.Q., Zhao R.L., Hyde K.D., Begerow D., Kemler M., Yurkov A., McKenzie E.H.C., Raspé O., Makoto K., Sánchez-Ramírez S., Vellinga E.C., Halling R., Papp V., Zmitrovich I.V., Buyck B., Ertz D., Wijayawardene N.N., Cui B.K., Schoutteten N., Liu X.Z., Li T.H., Yao Y.J., Zhu X.Y., Liu A.Q., Li G.J., Zhang M.Z., Ling Z.L., Cao B., Antonín V., Boekhout T., Barbosa da Silva B.D., De Crop E., Decock C., Dima B., Dutta A.K., Fell J.W., Geml J., Ghobad-Nejhad M., Giachini A.J., Gibertoni T.B., Gorjón S.P., Haelewaters D., He S.H., Hodkinson B.P., Horak E., Hoshino T., Justo A., Lim Y.W., Menolli Jr. N., Mešić A., Moncalvo J.-M., Mueller G.M., Nagy L.G., Nilsson R.H., Noordeloos M., Nuytinck J., Orihara T., Ratchadawan C., Rajchenberg M., Silva-Filho A.G.S., Sulzbacher M.A., Tkalčec Z., Valenzuela R., Verbeken A., Vizzini A., Wartchow F., Wei T.Z., Weiß M., Zhao C.L., Kirk P.M., 2020. Notes, outline and divergence times of Basidiomycota. Fungal Diversity 99 (1): 105–367. doi:10.1007/s13225-019-00435-4

Camacho J., Heyde A., Bhullar B.A.S., Haelewaters D., Simmons N.B., Abzhanov A., 2019: Peramorphosis, an evolutionary developmental mechanism in neotropical bat skull diversity. Developmental Dynamics 248 (11): 1129–1143. doi:10.1002/dvdy.90

Johnston P.R, Quijada L., Smith C.A., Baral H.-O., Hosoya T., Baschien C., Pärtel K., Zhuang W.-Y., Haelewaters D., Park D., López-Giráldez F., Wang Z., Townsend J.P. 2019. A multigene phylogeny toward a new phylogenetic classification for the Leotiomycetes. IMA Fungus 10: 1.

Fryar S.C., Haelewaters D., Catcheside D.E.A. 2019. Annabella australiensis gen. & sp. nov. (Helotiales, Cordieritidaceae) from South Australian mangroves. Mycological Progress 18 (7): 973-981.

Saba M., Haelewaters D., Fiaz M., Khalid A.N., Pfister DH. 2019. Amanita mansehraensis, a new species in section Vaginatae from Pakistan. Phytotaxa 409 (4): 189-201.

Saba M., Haelewaters D., Iturriaga T., Khalid A.N., Ashraf T., Pfister D.H. 2019. Geopora ahmadii sp. nov. (Pyronemataceae, Pezizales), a new species from Pakistan. Mycotaxon 134 (2): 377-389.

Haelewaters D., Hiller T., Pan F.Y., Pan J.Y. 2019: Tracking an ectoparasitic fungus of Harmonia axyridis in the USA using literature records and citizen science data. IOBC-WPRS Bull. 145: 16–21.

Haelewaters D. 2019: Hesperomyces “harmoniae” nom. prov. (Laboulbeniales), an ectoparasitic fungus specific to Harmonia axyridis. IOBC-WPRS Bulletin 145: 52–54.

Haelewaters D., Boer P., Báthori F., Rádai Z., Reboleira A.S.P.S., Tartally A., De Kesel A., Pfliegler W.P., Nedvěd O., 2019: Studies of Laboulbeniales on Myrmica ants (IV): host-related diversity and thallus distribution patterns of Rickia wasmannii. Parasite 26: 29. doi:10.1051/parasite/2019028

Hiller T., Haelewaters D. 2019. A case of silent invasion: citizen science confirms the presence of Harmonia axyridis (Coleoptera, Coccinellidae) in Central America. PLoS ONE 14(7): e0220082.

Szentiványi T., Haelewaters D., Rádai Z., Mizsei E., Pfliegler W.P., Báthori F., Tartally A., Glaizot O., Christe P., 2019: Climatic effects on the distribution of ant- and bat fly-associated fungal ectoparasites (Ascomycota, Laboulbeniales). Fungal Biol. 39: 371–379. doi:10.1016/j.funeco.2019.03.003

Biranvand A., Nedvěd O., Tomaszewska W., Al Ansi A.N., Fekrat L., Haghghadam Z.M., Khormizi M.Z., Noorinahad S., Şenal D., Shakarami J., Haelewaters D., 2019: The genus Harmonia (Coleoptera, Coccinellidae) in the Middle East. Acta Entomol. Mus. Nat. Prague 59 (1): 163–170. doi:10.2478/aemnp-2019-0014

De Kesel A., Haelewaters D., 2019: Laboulbeniales (Fungi, Ascomycota) of cholevine beetles (Coleoptera, Leiodidae) in Belgium and the Netherlands. Sterbeeckia 35: 60–66.

Haelewaters D., Pfister D.H., 2019: Morphological species of Gloeandromyces (Ascomycota, Laboulbeniales) evaluated using single-locus species delimitation methods. Fungal Syst. Evol. 3: 19–33. doi:10.3114/fuse.2019.03.03

Haelewaters D., Pfliegler W.P., Gorczak M., Pfister D.H., 2019: Birth of an order: comprehensive molecular phylogenetic study reveals that Herpomyces (Fungi, Laboulbeniomycetes) is not part of Laboulbeniales. Mol. Phylogenet. Evol. 133: 286–301. doi:10.1016/j.ympev.2019.01.007

Haelewaters D., Shapiro-Ilan D.I., Cottrell T.E., 2018: Will dual fungal infections increase mortality of Harmonia axyridis in natural populations? IOBC-WPRS Bull. 137: 13–17.

Haelewaters D., De Kesel A., Pfister D.H., 2018: Integrative taxonomy reveals hidden species within a common fungal parasite of ladybirds. Sci. Rep. 8 (1): 15966. doi:10.1038/s41598-018-34319-5

Haelewaters D., Hiller T., Dick C.W., 2018: Bats, bat flies, and fungi: a case of hyperparasitism. Trends Parasitol. 34 (9): 784–799. Doi:10.1016/


Conference presentations (selection):

Haelewaters D., Martin T.E. Bat fly-associated fungi: current developments and a call for global collaborations. Oral presentation, 49th North American Symposium on Bat Research, Kalamazoo MI. 26 October 2019.
– Haelewaters D. Patterns of speciation in thallus-forming Laboulbeniomycetes. Oral presentation, 18th Congress of European Mycologists, Warsaw-Białowieża (Poland). 16–21 September 2019.
– Szentiványi T., Pfliegler W.P., Sándor A.D., Christe P., Glaizot O., Blackwell M., Haelewaters D. Distribution, host specificity, and variation in Arthrorhynchus nycteribiae and A. eucampsipodae (Laboulbeniomycetes), an update. Poster presentation, 18th International Bat Research Conference. Phuket, Thailand. 28 July–1 August 2019.
Haelewaters D., Pan F.Y., Pan J.Y. Tracking an ectoparasitic fungus of Harmonia axyridis in North America using literature records and citizen science data. Poster presentation, IOBC-WPRS Working Group Meeting, Ponta Delgada, Azores. 13 September 2018.
Haelewaters D. Hesperomyces “harmoniae” nom. prov. (Laboulbeniales), an ectoparasitic fungus specific to Harmonia axyridis. Oral presentation, IOBC-WPRS Working Group Meeting, Ponta Delgada, Azores. 12 September 2018.
Haelewaters D., Pfister D.H. Herpomyces (Laboulbeniomycetes): a new order and a new species spread through pet trade. Poster presentation, 11th International Mycological Congress, San Juan PR, 17 July 2018.
Haelewaters D., Hiller T. Bats, bat flies, and Laboulbeniales fungi from the Chucantí Nature Reserve in eastern Panama. Oral presentation, 47th North American Symposium on Bat Research, Knoxville TN, 19 October 2017.
Haelewaters D. Murciélagos, moscas en murciélagos y hongos Laboulbeniales: un modelo para estudiar hiperparasitismo. Oral presentation, III Congreso Científico, Universidad Autonóma de Chiriquí, Davíd (Panama). 15 August 2017.
Haelewaters D., Pfliegler W.P., Gorczak M., Pfister D.H. Laboulbeniomycetes fungal ectoparasites of cockroaches: three-gene phylogeny and implications. Oral presentation, Evolution Annual Meeting, Portland OR. 23-27 June 2017.
– Pfliegler W.P., Haelewaters D., Báthori F., Zimmerman C.-U., Tartally A. Rickia wasmannii, a fungal ectoparasite of Myrmica ants, is a single phylogenetic species based on ribosomal DNA. Poster presentation, 7th Central European Workshop of Myrmecology, Krakow (Poland). 21-24 April 2017.
Haelewaters D. Can Laboulbeniales parasites (Fungi, Ascomycota) be invasive? Oral presentation, IOBC-WPRS Working Group Meeting, Bornholm, Denmark. 14 May 2015.
Haelewaters D., Shapiro-Ilan D.I., Cottrell T.E. Will dual fungal infections increase Harmonia axyridis mortality in natural populations? Oral presentation, IOBC-WPRS Working Group Meeting, Bornholm, Denmark. 13 May 2015.

Invited lectures (selection):

– Factors driving speciation in a group of fungal epibionts on arthropods. EcoLunch Seminar Series, Department of Biological Sciences, Purdue University, West Lafayette IN. 28 August 2019.
– Mycologist goes mammalogy: Laboulbeniales fungi on bat-associated flies. Invited talk, Cusuco National Park (Honduras). 13 July 2019.
– Taxonomy and species delimitation in Laboulbeniales (Fungi, Ascomycota). Invited talk, Ximena Bernal Lab Meeting, Department of Biological Sciences, Purdue University, West Lafayette IN. 24 April 2019.
– Laboulbeniales, their ecology, diversity and taxonomy. Invited lecture, Faculty of Science, Charles University, Czech Republic. 24-25 October 2018.
– Speciation and diversity in Laboulbeniomycetes. Invited seminar, Department of Botany and Plant Pathology, Purdue University, West Lafayette, Indiana, USA. 27 April 2018.
– On the hunt for fungi on flies on bats: A summer of fieldwork in Panama. Herbaria Seminar Series, Harvard University Herbaria, Cambridge, Massachusetts, USA. 26 September 2017.
– Three independent lineages of ectoparasitic fungi associated with bat flies. Invited seminar, Department of Zoology, University of South Bohemia, České Budějovice, Czech Republic. 23 August 2017.
– Fungal parasite on exotic lady beetle in South Africa: curse or blessing? [In Dutch]. Royal Academy for Overseas Sciences, Brussels, Belgium. 21 February 2017.
– Fungal ectoparasite spreading throughout the world in the footsteps of its novel main host, the globally invasive harlequin ladybird Harmonia axyridis. XXV International Congress of Entomology, Orlando, Florida, USA. 30 September 2016.
– Bats, bat flies, and Laboulbeniales fungi: a model for studying hyperparasitism. Bambi Seminar Series, Smithsonian Tropical Research Institute, Isla Barro Colorado, Panama. 14 July 2016.

Attended conferences:

– 2019: Mycological Society of America Annual Meeting, Minneapolis, Minnesota, USA
– 2018: IOBC-WPRS Working Group Meeting, Ponta Delgada, Azores
– 2018: 11th International Mycological Congress, San Juan, Puerto Rico
– 2017: 47th North American Symposium on Bat Research, Knoxville, Tennessee, USA
– 2017: III Congreso Científico, Universidad Autonóma de Chiriquí, Davíd, Panama
– 2017: Boston Harbor & Islands Science Symposium, Northeastern University, Boston, Massachusetts, USA
– 2016: XXV International Congress of Entomology, Orlando, Florida, USA
– 2016: Mycological Society of America Annual Meeting, Berkeley, California, USA
– 2016: Northeast Natural History Conference, Springfield, Massachusetts, USA
– 2015: Joint International Conference “Botany 2015”, Edmonton, Alberta, Canada
– 2015: New England Botanical Club 120th Anniversary Research Conference, Smith College, Northampton, Massachusetts, USA
– 2015: IOBC-WPRS Working Group Meeting, Bornholm, Denmark
– 2015: Second International Workshop on Ascomycete Systematics, Amsterdam, The Netherlands
– 2014: Mycological Society of America Annual Meeting, East Lansing, Michigan, USA
– 2013: American Phytopathological Society and Mycological Society of America Joint Meeting, Austin, Texas, USA

Membership in organizations:

– Indiana Academy of Science
– Koninklijke Vlaamse Mycologische Vereniging
– Mycological Society of America
– North American Society for Bat Research
– Society for the Study of Evolution

Back To Top